ICE SEAL PUBLICATIONS

This list is a work in progress. Please contact the Ice Seal Committee if you would like to add publications to the list.

2026

Lindsay, J. M., Conn, P. B., Boveng, P. L., Crawford, J. A., Quakenbush, L. T., Von Duyke, A. L., & Laidre, K. L. (2026). Ringed Seal (Pusa hispida) Haul-Out Behavior and Emergence Timing in the Bering, Chukchi, and Beaufort Seas. Ecology and Evolution, 16(1), e72948. https://doi.org/10.1002/ece3.72948

2025

Boveng, P.L., Chernook, V.I., Moreland, E.E., Conn, P.B., Trukhanova, I.S., Cameron, M.F., Christman, C.L., Crawford, J.A., Glazov, D., Harwood, L., Hou, B.X., Koslovsky, S.M., Lindsay, J.M., Litovka, D.I., London, J.M., McClintock, B.T., Platonov, N., Quakenbush, L., Richmond, E.L., Vasiliev, A., Von Duyke, A.L., Willoughby, A. (2025) Abundance and distribution of ringed and bearded seals in the Chukchi Sea: A reference for future trends. Arctic Science. https://doi.org/10.1139/as-2024-0068

Cameron, M., Brady, G., Hou, B., Koslovsky, S., McClintock, B., Meranda, M. Reichmuth, C., Richmond, E., and Ziel, H. (2025) Final report for the NOAA ORTA Bridging Program Project: an Extension of 2020-05 monitoring body condition of seals in Alaska using small UAS. AFSC Processed Rep. 2025-03, 46 p. Alaska Fish. Sci. Cent., NOAA, Natl. Mar. Fish. Serv., 7600 Sand Point Way NE, Seattle WA 98115. https://repository.library.noaa.gov/view/noaa/69885

Gryba, R., Von Duyke, A., Huntington, H. P., Adams, B., Frantz, B., Gatten, J., Harcharek, Q., Sarren, R., Henry, G., & Auger-Méthé, M. (2025). Indigenous Knowledge as a sole data source in habitat selection functions. Proceedings of the National Academy of Sciences, 122(15), e2411946122. https://doi.org/10.1073/pnas.2411946122

McCarthy, M. L., Martínez, A. R., Ferguson, S. H., Rosing-Asvid, A., Dietz, R., De Cahsan, B., Schreiber, L., Lorenzen, E. D., Hansen, R. G., Stimmelmayr, R., Bryan, A., Quakenbush, L., Lydersen, C., Kovacs, K. M., & Olsen, M. T. (2025). Circumpolar Population Structure, Diversity and Recent Evolutionary History of the Bearded Seal in Relation to Past and Present Icescapes. Molecular Ecology, n/a(n/a), e17643. https://doi.org/10.1111/mec.1764

Olnes, J., Bryan, A. L., Stimmelmayr, R., & Quakenbush, L. (2025). Organochlorine concentrations in ice seals harvested in Alaska in the 2000s and 2010s. Marine Pollution Bulletin, 215, 117843. https://doi.org/10.1016/j.marpolbul.2025.117843

Olnes, J., Quakenbush, L., Bryan, A., & Stimmelmayr, R. (2025). Trace elements in Alaska’s ice seals in the 2000s and 2010s. Science of The Total Environment, 958, 178126. https://doi.org/10.1016/j.scitotenv.2024.178126

Packard, N., Sills, J. M., Jones, R. A., Aldana, S. T., & Reichmuth, C. (2025). Sensitivity to airborne sounds in ice-dependent bearded seals. Journal of Comparative Physiology A. https://doi.org/10.1007/s00359-025-01782-1

Regehr, E. V., Quakenbush, L., Von Duyke, A. L., Citta, J. J., & Lindsay, J. M. (2025). Projecting the future of a threatened marine mammal in relation to climate warming. Ecological Applications, 35(8), e70127. https://doi.org/10.1002/eap.70127

Reichmuth, C., and Klein, A.M. Bearded Seal Erignathus barbatus (Erxleben, 1777). (2025) In: Hackländer, K., Zachos, F.E. (eds) Handbook of the Mammals of Europe. Handbook of the Mammals of Europe. Springer, Cham. https://doi.org/10.1007/978-3-319-65038-8_141-1

Sills, J. M., Ruscher, B., Packard, N., Jones, R. A., Southall, B. L., and Reichmuth, C. (2025) Low-frequency hearing and masking parameters in representative seals and sea lions. The Journal of the Acoustical Society of America 158(2), 1585-1599. https://doi.org/10.1121/10.0039049


2024

Ferrer, T., Boveng, P., Hauser, D. D. W., Withrow, D., Burkanov, V., Quinn, T. P., & O’Corry-Crowe, G. (2024). Genetic and evolutionary divergence of harbour seals (Phoca vitulina) in Iliamna Lake, Alaska. Biology Letters, 20(10), 20240166. https://doi.org/10.1098/rsbl.2024.0166

Florko, K. R. N., Yurkowski, D. J., Breiter, C.-J. C., Ferguson, S. H., Gamblin, H. E. L., Grottoli, A., Nace, A., Shuert, C. R., & Petersen, S. D. (2024). Biologging reveals rapid movements of harbour seals between freshwater and marine habitats in the subarctic. Frontiers in Marine Science, 11. https://www.frontiersin.org/journals/marine-science/articles/10.3389/fmars.2024.1435206

Frouin-Mouy, H., Rountree, R., Juanes, F., Aguzzi, J., & De Leo, F. C. (2024). Deep-sea cabled video-observatory provides insights into the behavior at depth of sub-adult male northern elephant seals, Mirounga angustirostris. PLOS ONE, 19(9), e0308461. https://doi.org/10.1371/journal.pone.0308461

Gryba, R. (2024). New approaches to understand species-habitat relationship using Indigenous Knowledge and scientific data [PhD Dissertation, University of British Columbia]. https://dx.doi.org/10.14288/1.0440972

London, J. M., Conn, P. B., Koslovsky, S. M., Richmond, E. L., Ver Hoef, J. M., Cameron, M. F., Crawford, J. A., Von Duyke, A. L., Quakenbush, L., & Boveng, P. L. (2024). Spring haul-out behavior of seals in the Bering and Chukchi Seas: Implications for abundance estimation. PeerJ, 12, e18160. https://doi.org/10.7717/peerj.18160

Morgenthal, K., Krüger, Y., Rogers, T., Dehnhardt, G., & Hanke, F. D. (2024). Characterization of pinniped vibrissal type and number. Marine Mammal Science, n/a(n/a), e13166. https://doi.org/10.1111/mms.13166

Pace, C., Goertz, C. E. C., Hunter, N., Abraham, T., Goertz, J., & Reichmuth, C. (2024). Seasonal variation of blood analytes associated with skin health in Alaskan ice seals. Journal of Zoo and Wildlife Medicine, 55(4), 994–1004. https://doi.org/10.1638/2024-0046

Quakenbush, L., Bryan, A., Crawford, J., Olnes, J., & Stimmelmayr, R. (2024). NOAA Arctic Report Card 2024: Ice Seals of Alaska. NOAA Technical Report OAR ARC ; 24-12 (Arctic Report Card). (noaa:66868). https://doi.org/10.25923/4488-8843

Taras, B. D., Conn, P. B., Bravington, M. V., Kilian, A., Lang, A. R., Bryan, A., Stimmelmayr, R., & Quakenbush, L. (2024). Estimating Demographic Parameters for Bearded Seals, Erignathus barbatus, in Alaska Using Close-Kin Mark-Recapture Methods. Evolutionary Applications, 17(11), e70035. https://doi.org/10.1111/eva.70035

Tengler, M. L., Dearolf, J., Bryan, A. L., Reichmuth, C., and Thometz, N. M. (2024) Comparative Muscle Physiology of Ringed (Pusa hispida), Bearded (Erignathus barbatus), and Spotted (Phoca largha) Seals from the Bering and Chukchi Seas. Aquatic Mammals 50(3), 181-198. https://doi.org/10.1578/AM.50.3.2024.181


2023

de la Vega, C., Kershaw, J., Stenson, G. B., Frie, A. K., Biuw, M., Haug, T., Norman, L., Mahaffey, C., Smout, S., & Jeffreys, R. M. (2023). Multi-decadal trends in biomarkers in harp seal teeth from the North Atlantic reveal the influence of prey availability on seal trophic position. Global Change Biology, n/a(n/a). https://doi.org/10.1111/gcb.16889

Hoseini, S. M., Namroodi, S., Sayadshirazi, A., & Zaccaroni, A. (2023). Trace Elements and Contaminants Concentrations in Tissues of Caspian Seals (Pusa caspica) along the Iranian Coast. Toxics, 11(1). https://doi.org/10.3390/toxics11010039

Lindsay, J., Hauser, D., Mahoney, A., Laidre, K., Goodwin, J., Harris, C., Schaeffer, R., Schaeffer R, S., Whiting, A., Boveng, P., Laxague, N., Betcher, S., Subramaniam, A., Witte, C., & Zappa, C. (2023). Characteristics of ringed seal Pusa hispida (‘natchiq’) denning habitat in Kotzebue Sound, Alaska, during a year of limited sea ice and snow. Marine Ecology Progress Series, 705, 1–20. https://doi.org/10.3354/meps14252

Olnes, J., Crawford, J., Okkonen, S., Citta, J. J., Quakenbush, L., Von Duyke, A. L., Maslowski, W., Osinski, R., & Druckenmiller, M. L. (2023). Oceanographic influences on spotted seal foraging in the Pacific Arctic. Continental Shelf Research, 105136. https://doi.org/10.1016/j.csr.2023.105136

Rosing-Asvid, A., Löytynoja, A., Momigliano, P., Hansen, R. G., Scharff-Olsen, C. H., Valtonen, M., Kammonen, J., Dietz, R., Rigét, F. F., Ferguson, S. H., Lydersen, C., Kovacs, K. M., Holland, D. M., Jernvall, J., Auvinen, P., & Tange Olsen, M. (2023). An evolutionarily distinct ringed seal in the Ilulissat Icefjord. Molecular Ecology, 32(22), 5932–5943. https://doi.org/10.1111/mec.17163

Thometz, N. M., Rosen, D. A. S., Hermann-Sorensen, H., Meranda, M., Pardini, M., & Reichmuth, C. (2023). Maintaining control: Metabolism of molting Arctic seals in water and when hauled out. Journal of Experimental Biology, 226(1), jeb244862. https://doi.org/10.1242/jeb.244862


2022

Biard, V., Nykänen, M., Niemi, M., & Kunnasranta, M. (2022). Extreme moulting site fidelity of the Saimaa ringed seal. Mammalian Biology. https://doi.org/10.1007/s42991-021-00209-z

Niemi, M., Nykänen, M., Biard, V., Kurkilahti, M., & Kunnasranta, M. (2022). Molting phenology of a lacustrine ringed seal, Pusa hispida saimensis. Ecology and Evolution, 12(8), e9248. https://doi.org/10.1002/ece3.9248

Olnes, J., Quakenbush, L., Nelson, M., Simon, A., Burns, J., and the Ice Seal Committee (2022) Trends in subsistence harvests of ice seals in the Yukon-Kuskokwim Delta region, Alaska, 1962-2018. Arctic 75(4):449-461. https://doi.org/10.14430/arctic76302

Scherdin, N., Djukarić, J., & Desportes, G. (2022). Bearded seals in the Atlantic Arctic: Review of post 2010 knowledge available for informing stock assessments. NAMMCO Scientific Publications, 12. https://doi.org/10.7557/3.6883

Sills, J. M., & Reichmuth, C. (2022). Vocal Behavior in Spotted Seals (Phoca largha) and Implications for Passive Acoustic Monitoring. Frontiers in Remote Sensing, 3. https://doi.org/10.3389/frsen.2022.862435


2021

Andersen, M., Kovacs, K. M., & Lydersen, C. (2021). Stable ringed seal (Pusa hispida) demography despite significant habitat change in Svalbard, Norway. Polar Research, 40. https://doi.org/10.33265/polar.v40.5391

Crain, D.D., Karpovich, S.A., Quakenbush, L., Polasek, L. (2021) Using claws to compare reproduction, stress and diet of female bearded and ringed seals in the Bering and Chukchi seas, Alaska, between 1953–1968 and 1998–2014. Conservation Physiology 9(1):coaa115.  https://doi.org/10.1093/conphys/coaa115

Fournet, M. E. H., Silvestri, M., Clark, C. W., Klinck, H., & Rice, A. N. (2021). Limited vocal compensation for elevated ambient noise in bearded seals: Implications for an industrializing Arctic Ocean. Proceedings of the Royal Society B: Biological Sciences, 288(1945), 20202712. https://doi.org/10.1098/rspb.2020.2712

Gryba, R., Huntington, H. P., Von Duyke, A. L., Adams, B., Frantz, B., Gatten, J., Harcharek, Q., Olemaun, H., Sarren, R., Skin, J., Henry, G., & Auger-Méthé, M. (2021). Indigenous Knowledge of bearded seal (Erignathus barbatus), ringed seal (Pusa hispida), and spotted seal (Phoca largha) behaviour and habitat use near Utqiaġvik, Alaska, USA. Arctic Science, 7, 832–858. https://doi.org/10.1139/as-2020-0052

Hauser, D. D. W., Frost, K. J., & Burns, J. J. (2021). Ringed seal (Pusa hispida) breeding habitat on the landfast ice in northwest Alaska during spring 1983 and 1984. PLOS ONE, 16(11), e0260644. https://doi.org/10.1371/journal.pone.0260644

Hendrix, A. M., Lefebvre, K. A., Quakenbush, L., Bryan, A., Stimmelmayr, R., Sheffiled, G., Wosswaesser, G., Willis, M. L., Bowers, E. K., Kendrick, P., Frame, E., Burbacher, T., & Marcinek, D. J. (2021). Ice seals as sentinels for algal toxin presence in the Pacific Arctic and subarctic marine ecosystems. Marine Mammal Science, 1–17. https://doi.org/10.1111/mms.12822

Kovacs, K. M., Citta, J., Brown, T., Dietz, R., Ferguson, S., Harwood, L., Houde, M., Lea, E. V., Quakenbush, L., Riget, F., Rosing-Asvid, A., Smith, T., Svetochev, V., Svetocheva, O., & Lydersen, C. (2021). Variation in body size of ringed seals (Pusa hispida hispida) across the circumpolar Arctic: Evidence of morphs, ecotypes or simply extreme plasticity? Polar Research, 40. https://doi.org/10.33265/polar.v40.5753

Ogloff, W. R., Ferguson, S. H., Fisk, A. T., Marcoux, M., Hussey, N. E., Jaworenko, A., & Yurkowski, D. J. (2021). Long-distance movements and associated diving behaviour of ringed seals (Pusa hispida) in the eastern Canadian Arctic. Arctic Science, 7(2), 494–511. https://doi.org/10.1139/as-2019-0042

Rode, K.D., Regehr, E.V., Bromaghin, J.F., Wilson, R.R., St. Martin, M., Crawford, J.A., and Quakenbush, L.T. (2021). Seal body condition and atmospheric circulation patterns influence polar bear body condition, recruitment, and feeding ecology in the Chukchi Sea.  Global Change Biology 27(12): 2684-2701. https://doi.org/10.1111/gcb.15572

Thometz, N. M., Hermann-Sorensen, H., Russell, B., Rosen, D. A. S., & Reichmuth, C. (2021). Molting strategies of Arctic seals drive annual patterns in metabolism. Conservation Physiology, 9(1), coaa112. https://doi.org/10.1093/conphys/coaa112


2020

Davidson, S., et al. (2020).  New ecological insights from the Arctic Animal Movement Archive (AAMA).  Science.  Science 370(6517):712-715. https://doi.org/10.1126/science.abb7080

Kennedy, S., Kuiken, T., Jepson, P. D., Deaville, R., Forsyth, M., Barrett, T., van de Buildt, M. W. G., Osterhaus, A. D. M. E., Eybatov, T., Duck, C., Kydyrmanov, A., Mitrofanov, I., & Wilson, S. (2000). Mass die-off of Caspian seals caused by canine distemper virus. Emerging Infectious Diseases, 6(6), 637–645. https://doi.org/10.3201/eid0606.000613

Olnes, J., Breed, G., Druckenmiller, M., Citta, J.J., Crawford, J.A., Von Duyke, A.L., Frost, K.J., and Quakenbush, L. (2020). Juvenile bearded seal response to a decade of sea ice change in the Bering, Chukchi, and Beaufort seas. Marine Ecology Progress Series.  Marine Ecology Progress Series. https://doi.org/10.3354/meps13609
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Olnes, J., Crawford, J., Citta, J. J., Druckenmiller, M. L., Von Duyke, A. L., & Quakenbush, L. (2020). Movement, diving, and haul-out behaviors of juvenile bearded seals in the Bering, Chukchi and Beaufort seas, 2014–2018. Polar Biology. https://doi.org/10.1007/s00300-020-02710-6

Schmidt, B., Sonne, C., Nachtsheim, D., Dietz, R., Oheim, R., Rolvien, T., Persson, S., Amling, M., & Siebert, U. (2020). Variation in skull bone mineral density of ringed seals (Phoca hispida) from the Gulf of Bothnia and West Greenland between 1829 and 2019. Environment International, 143, 105968. https://doi.org/10.1016/j.envint.2020.105968

Sills, J. M., Reichmuth, C., Southall, B. L., Whiting, A., & Goodwin, J. (2020). Auditory biology of bearded seals (Erignathus barbatus). Polar Biology. https://doi.org/10.1007/s00300-020-02736-w

Von Duyke, A. L., Douglas, D. C., Herreman, J. K., & Crawford, J. A. (2020). Ringed seal (Pusa hispida) seasonal movements, diving, and haul-out behavior in the Beaufort, Chukchi, and Bering Seas (2011–2017). Ecology and Evolution, 10(12), 5595–5616. https://doi.org/10.1002/ece3.6302

Walden, H.S., Bryan, A.L., McIntosh, A., Tuomi, P., Hoover-Miller, A., Stimmelmayr, R., and Quakenbush, L. (2020). Helminth fauna of ice seals in the Alaskan Bering and Chukchi Seas, 2006-15. Journal of Wildlife Diseases 56(4):863-872.  https://doi.org/10.7589/2019-09-228

Yurkowski, D. J., Richardson, E. S., Lunn, N. J., Muir, D. C. G., Johnson, A. C., Derocher, A. E., Ehrman, A. D., Houde, M., Young, B. G., Debets, C. D., Sciullo, L., Thiemann, G. W., & Ferguson, S. H. (2020). Contrasting Temporal Patterns of Mercury, Niche Dynamics, and Body Fat Indices of Polar Bears and Ringed Seals in a Melting Icescape. Environmental Science & Technology, 54(5), 2780–2789. https://doi.org/10.1021/acs.est.9b06656


2019

Goertz, C.E.C., Reichmuth, C., Thometz, N.M., Ziel, H., and Boveng, P. (2019) Comparative Health Assessments of Alaskan Ice Seals. Frontiers in Veterinary Science 6(4):1-15.  https://doi.org/10.3389/fvets.2019.00004

Gryba, R., Wiese, F., Kelly, B., Von Duyke, A., Pickart, R., & Stockwell, D. (2019). Inferring foraging locations and water masses preferred by spotted seals Phoca largha and bearded seals Erignathus barbatus. Marine Ecology Progress Series, 631, 209–224. https://doi.org/10.3354/meps13145

Nelson, M. A., Quakenbush, L. T., Taras, B. D., & Ice Seal Committee. (2019). Subsistence harvest of ringed, bearded, spotted, and ribbon seals in Alaska is sustainable. Endangered Species Research, 40, 1–16. https://doi.org/10.3354/esr00973

Reimer, J. R., Brown, H., Beltaos-Kerr, E., & de Vries, G. (2019). Evidence of intraspecific prey switching: Stage-structured predation of polar bears on ringed seals. Oecologia, 189(1), 133–148. https://doi.org/10.1007/s00442-018-4297-x

Reimer, J. R., Caswell, H., Derocher, A. E., & Lewis, M. A. (2019). Ringed seal demography in a changing climate. Ecological Applications, 29(3), e01855. https://doi.org/10.1002/eap.1855


2018

Cameron, M. F., Frost, K. J., Ver Hoef, J. M., Breed, G. A., Whiting, A. V., Goodwin, J., & Boveng, P. L. (2018). Habitat selection and seasonal movements of young bearded seals (Erignathus barbatus) in the Bering Sea. PLOS ONE, 13(2), e0192743. https://doi.org/10.1371/journal.pone.0192743

Citta, J., Lowry, L., Quakenbush, L., Kelly, B., Fischbach, A., London, J., Jay, C., Frost, K., O’Corry Crowe, G., Crawford, J., Boveng, P., Cameron, M., Von Duyke, A., Nelson, M., Harwood, L., Richard, P., Suydam, R., Heide-Jorgensen, M.P., Hobbs, R., Litovka, D., Whiting, A., Marcoux, M., Kennedy, A., George, J., Orr, J., Gray, T. (2018). A multi-species synthesis of satellite telemetry data in the Pacific Arctic (1987–2015): Overlap of marine mammal distributions and core use areas. Deep Sea Research II 152:132-153. https://doi.org/10.1016/j.dsr2.2018.02.006

Crawford, J.A., Frost, K.J., Quakenbush, L.T., and Whiting, A. (2018) Seasonal and diel differences in dive and haul-out behavior of adult and subadult ringed seals (Pusa hispida) in the Bering and Chukchi seas. Polar Biology 41:65–80. https://doi.org/10.1007/s0030 0-018-2399-x

Frost, K. J., & Burns, J. J. (2018). Spotted Seal: Phoca largha. In B. Würsig, J. G. M. Thewissen, & K. M. Kovacs (Eds.), Encyclopedia of Marine Mammals (Third Edition) (pp. 928–931). Academic Press. https://doi.org/10.1016/B978-0-12-804327-1.00244-2

Jensen, A. M., Sheehan, G. W., & MacLean, S. A. (2018). Inuit and Marine Mammals. In B. Würsig, J. G. M. Thewissen, & K. M. Kovacs (Eds.), Encyclopedia of Marine Mammals (Third Edition) (pp. 520–525). Academic Press. https://doi.org/10.1016/B978-0-12-804327-1.00160-6


2017

Boveng, P. L., Cameron, M. F., Conn, P. B., & Moreland, E. E. (2017). Abundance Estimates of Ice-Associated Seals: Bering Sea Populations that Inhabit the Chukchi Sea During the Open-Water Period (BOEM Report 2016-077; p. 119 with appendices). https://espis.boem.gov/final%20reports/5578.pdf?utm_source=chatgpt.com

Lydersen, C., Vaquie-Garcia, J., Lydersen, E., Christensen, G. N., & Kovacs, K. M. (2017). Novel terrestrial haul-out behaviour by ringed seals (Pusa hispida) in Svalbard, in association with harbour seals (Phoca vitulina). Polar Research, 36(0). https://polarresearch.net/index.php/polar/article/view/2658

Wilson, S. C., Dolgova, E., Trukhanova, I., Dmitrieva, L., Crawford, I., Baimukanov, M., & Goodman, S. J. (2017). Breeding behavior and pup development of the Caspian seal, Pusa caspica. Journal of Mammalogy, 98(1), 143–153. https://doi.org/10.1093/jmammal/gyw176

2016

Huntington, H.P., Quakenbush, L.T., and Nelson, M. (2016). Effects of changing sea-ice on marine mammals and subsistence hunters in northern Alaska from traditional knowledge interviews. Biology Letters 12:20160198. https://doi.org/10.1098/rsbl.2016.0198

2015

Crawford, J.A., Quakenbush, L.T., and Citta, J.J. (2015). A comparison of ringed and bearded seal diet, condition and productivity between historical (1975–1984) and recent (2003–2012) periods in the Alaskan Bering and Chukchi seas. Progress in Oceanography 136:133–150. https://doi.org/10.1016/j.pocean.2015.05.011

Harwood, L.A., Smith, T.G., Auld, J.C., Melling, H., and Yurkowski, D.J. (2015). Seasonal movements and diving of ringed seals, Pusa hispida, in the western Canadian Arctic, 199–2001 and 2010–11. Arctic 68:193–209. https://doi.org/10.14430 /arcti c4479

​Sills, J.M., Southall, B.L., and Reichmuth, C. (2015). Amphibious hearing in ringed seals (Pusa hispida): underwater audiograms, aerial audiograms and critical ratio measurements.  Journal of Experimental biology 218:2250-2059.  https://doi.org/10.1242/jeb.120972

2014

Sills, J. M., Southall, B. L., & Reichmuth, C. (2014). Amphibious hearing in spotted seals (Phoca largha): Underwater audiograms, aerial audiograms and critical ratio measurements. The Journal of Experimental Biology, 217(5), 726. https://doi.org/10.1242/jeb.097469

2013

Moreland, E. E., Cameron, M., & Boveng, P. (2013). Bering Okhotsk Seal Surveys (BOSS) Joint U.S.-Russian Aerial Surveys for Ice-associated Seals, 2012-13 (Quarterly July August September; p. 6). NOAA AFSC. https://apps-afsc.fisheries.noaa.gov/Quarterly/jas2013/JAS13-Feature.pdf


2012

Crawford, J. A., Frost, K. J., Quakenbush, L. T., & Whiting, A. (2012). Different habitat use strategies by subadult and adult ringed seals (Phoca hispida) in the Bering and Chukchi seas. Polar Biology, 35(2), 241–255. https://doi.org/10.1007/s00300-011-1067-1

Harwood, L.A., Smith, T.G., and Auld, J.C. (2012) Fall migration of ringed seals (Phoca hispida) through the Beaufort and Chukchi Seas, 2001–02. Arctic 65:35–44. https://doi.org/10.14430 /arctic4163

Pilfold, N. W., Derocher, A. E., Stirling, I., Richardson, E., & Andriashek, D. (2012). Age and Sex Composition of Seals Killed by Polar Bears in the Eastern Beaufort Sea. PLOS ONE, 7(7), e41429. https://doi.org/10.1371/journal.pone.0041429


2010

Cameron, M. F., Bengtson, J. L., Boveng, P. L., Jansen, J. K., Kelly, B. P., Dahle, S. P., Logerwell, E. A., Overland, J. E., Sabine, C. L., Waring, G. T., & Wilder, J. M. (2010). Status Review of the Bearded Seal (Erignathus barbatus). https://repository.library.noaa.gov/view/noaa/3761

Kelly, B. P., Badajos, O. H., Kunnasranta, M., Moran, J. R., Martinez-Bakker, M., Wartzok, D., & Boveng, P. (2010). Seasonal home ranges and fidelity to breeding sites among ringed seals. Polar Biology, 33(8), 1095–1109. https://doi.org/10.1007/s00300-010-0796-x

Kelly, B. P., Bengtson, J. L., Boveng, P. L., Cameron, M. F., Dahle, S. P., Jansen, J. K., Logerwell, E. A., Overland, J. E., Sabine, C. L., Waring, G. T., & Wilder, J. M. (2010). Status Review of the Ringed Seal (Phoca hispida). https://apps-afsc.fisheries.noaa.gov/Publications/AFSC-TM/NOAA-TM-AFSC-212.pdf


2009

Boveng, P. L., Bengtson, J. L., Buckley, T. W., Cameron, M. F., Dahle, S. P., Kelly, B. P., Megrey, B. A., Overland, J. E., & Williamson, N. J. (2009). Status Review of the Spotted Seal (Phoca largha). https://repository.library.noaa.gov/view/noaa/3671

Bowen, W. D., Beck, C. A., & Austin, D. A. (2009). Pinniped Ecology. In W. F. Perrin, B. Würsig, & J. G. M. Thewissen (Eds.), Encyclopedia of Marine Mammals (Second Edition) (pp. 852–861). Academic Press. https://doi.org/10.1016/B978-0-12-373553-9.00198-X

Quakenbush, L., Citta, J., & Crawford, J. (2009). Biology of the Spotted Seal (Phoca largha) in Alaska from 1962 to 2008. Report to: National Marine Fisheries Service. Alaska Department of Fish & Game - Arctic Marine Mammals Program. https://www.adfg.alaska.gov/static/research/programs/marinemammals/pdfs/biomonitoring_biology_spotted_seal_final_report.pdf


2008

Boveng, P.L., Bengtson, J.L., Buckley, T.W., Cameron, M.F., Dahle, S.P., Megrey, B.A., Overland, J.E., and Williamson, N.J. (2008). Status review of the ribbon seal (Histriophoca fasciata). U.S. Department of Commerce, NOAA Technical Memorandum NMFSAFSC-191. 115 p. https://www.fisheries.noaa.gov/resource/document/status-review-ribbon-seal-2008

Moore, S. E. (2008). Marine Mammals as Ecosystem Sentinels. Journal of Mammalogy, 89(3), 534–540. https://doi.org/10.1644/07-MAMM-S-312R1.1

Quakenbush, L. and Citta, J. (2008) Biology of the ribbon seal in Alaska. Report to National Marine Fisheries Service. 45 p. Arctic Marine Mammal Program, Alaska Department of Fish and Game, Fairbanks, AK. https://www.fisheries.noaa.gov/resource/peer-reviewed-research/biology-ribbon-seal-alaska


2006

Swanson, B. J., Kelly, B. P., Maddox, C. K., & Moran, J. R. (2006). Shed skin as a source of DNA for genotyping seals. Molecular Ecology Notes, 6(4), 1006–1009. https://doi.org/10.1111/j.1471-8286.2006.01473.x


2005

Bengtson, J. L., Hiruki-Raring, L. M., Simpkins, M. A., & Boveng, P. L. (2005). Ringed and bearded seal densities in the eastern Chukchi Sea, 1999–2000. Polar Biology, 28, 833–845. hard copy. https://doi.org/10.1007/s00300-005-0009-1

Bradford, A. L., & Weller, D. W. (2005). Spotted seal haul-out patterns in a coastal lagoon on Sakhalin Island, Russia. Mammal Study, 30(2), 145–149. https://doi.org/10.3106/1348-6160(2005)30%255B145:SSHPIA%255D2.0.CO;2

Moulton, V. D., Richardson, W. J., Elliott, R. E., McDonald, T. L., Nations, C., & Williams, M. T. (2005). Effects of an offshore oil development on local abundance and distribution of ringed seals (Phoca hispida) of the Alaskan Beaufort Sea. Marine Mammal Science, 21(2), 217–242. https://doi.org/10.1111/j.1748-7692.2005.tb01225.x

2004

Blackwell, S. B., Lawson, J. W., & Williams, M. T. (2004). Tolerance by ringed seals (Phoca hispida) to impact pipe-driving and construction sounds at an oil production island. Journal of the Acoustical Society of America, 115(5), 2346–2357. https://doi.org/10.1121/1.1701899

Frost, K. J., Lowry, L. F., Pendleton, G., & Nute, H. R. (2004). Factors Affecting the Observed Densities of Ringed Seals, Phoca hispida, in the Alaskan Beaufort Sea, 1996-99. Arctic, 57(2), 115–128. https://doi.org/10.14430/arctic489


2003

Boveng, P. L., Bengtson, John L., Withrow, D. E., Cesarone, J. C., Simpkins, M. A., Frost, K. J., & Burns, J. J. (2003). The abundance of harbor seals in the Gulf of Alaska. Marine Mammal Science, 19(1), 111–127. https://doi.org/10.1111/j.1748-7692.2003.tb01096.x

Moulton, V. D., Richardson, W. J., & Williams, M. T. (2003). Ringed seal densities and noise near an icebound artificial island with construction and drilling. Acoustics Research Letters Online, 4, 112. https://doi.org/10.1121/1.1605091

Ver Hoef, J. M., & Frost, K. J. (2003). A Bayesian hierarchical model for monitoring harbor seal changes in Prince William Sound, Alaska. Environmental and Ecological Statistics, 10, 201–219. https://doi.org/10.1023/A:1023626308538


2002

Moulton, V. D., Richardson, W. J., McDonald, T. L., Elliott, R. E., & Williams, M. T. (2002). Factors influencing local abundance and haulout behaviour of ringed seals (Phoca hispida) on landfast ice of the Alaskan Beaufort Sea. Canadian Journal of Zoology, 80, 1900–1917. https://doi.org/10.1139/z02-173

Trukhin, A. M., & Mizuno, A. W. (2002). Distribution and abundance of the largha seal (Phoca largha Pall.) on the coast of Primorye Region (Rusia): A literature review and survey report. Mammal Study, 27, 1–14. https://doi.org/10.3106/mammalstudy.27.1


2001

Frost, K. J., Simpkins, M. A., & Lowry, L. F. (2001). Diving behavior of subadult and adult harbor seals in Prince William Sound, Alaska. Marine Mammal Science, 17(4), 813–834. https://doi.org/10.1111/j.1748-7692.2001.tb01300.x

Lowry, L. F., Frost, K. J., Ver Hoef, J. M., & DeLong, R. A. (2001). Movements of satellite-tagged subadult and adult harbor seals in Prince William Sound, Alaska. Marine Mammal Science, 17(4), 835–861. https://doi.org/10.1111/j.1748-7692.2001.tb01301.x

Woshner, V. M., O’Hara, T. M., Bratton, G. R., & Beasley, V. R. (2001). Concentrations and interactions of selected essential and non-essential elements in ringed seals and polar bears of arctic Alaska. Journal of Wildlife Diseases, 37(4), 711–721. https://doi.org/10.7589/0090-3558-37.4.711


2000

Lowry, L. F., Burkanov, V. N., Frost, K. J., Simpkins, M. A., Davis, R., DeMaster, D. P., Suydam, R., & Springer, A. (2000). Habitat use and habitat selection by spotted seals (Phoca largha) in the Bering Sea. Canadian Journal of Zoology, 78(11), 1959–1971. https://doi.org/10.1139/z00-138


1999

Adams, A. M., & Rausch, R. L. (1989). A revision of the genus Orthosplanchnus Odhner, 1905 with consideration of the genera Odhneriella Skriabin, 1915 and Hadwenius Price, 1932 (Digenea: Campulidae). Canadian Journal of Zoology, 67, 1268–1278. https://doi.org/10.1139/z89-181

Cassini, M. H. (1999). The evolution of reproductive systems in pinnipeds. Behavioral Ecology, 10(5), 612–616. https://doi.org/10.1093/beheco/10.5.612

Frost, K. J., Lowry, L. F., & Ver Hoef, J. M. (1999). Monitoring the trend of harbor seals in Prince William Sound, Alaska, after the Exxon Valdez oil spill. Marine Mammal Science, 15(2), 494–506. https://doi.org/10.1111/j.1748-7692.1999.tb00815.x


1997

DeMaster, D. P., Rugh, D. J., Jones, B., Lowry, L. F., Frost, K. J., & Sheffield, G. (1997). Summaries of data collected from ice-associated seals in the Bering, Chukchi, and Beaufort Seas, 1975-1994.

Iverson, S. J., Frost, K. J., & Lowry, L. F. (1997). Fatty acid signatures reveal fine scale structure of foraging distribution of harbor seals and their prey in Prince William Sound, Alaska. Marine Ecology Progress Series, 151, 255–271. http://dx.doi.org/10.3354/meps151255

Rugh, D. J., Shelden, K. E. W., & Withrow, D. E. (1997). Spotted seals, Phoca largha, in Alaska. Marine Fisheries Review, 59(1), 1–18. Gale Academic OneFile. https://media.fisheries.noaa.gov/dam-migration/ak1997sesp-508.pdf


1996

Furgal, C. M., Innes, S., & Kovacs, K. M. (1996). Characteristics of ringed seal, Phoca hispida, subnivean structures and breeding habitat and their effects on predation. Canadian Journal of Zoology, 74, 858–874. https://doi.org/10.1139/z96-100

Kelly, B. P., & Wartzok, D. (1996). Ringed seal diving behavior in the breeding season. Canadian Journal of Zoology, 74, 1547–1555. https://doi.org/10.1139/z96-169

Lowry, L. F., Burkanov, V. N., & Frost, K. J. (1996). Importance of walleye pollock, Theragra chalcogramma, in the diet of phocid seals in the Bering Sea and northwestern Pacific Ocean. NOAA Technical Report NMFS 1996(126): 141-151. http://www.adfg.alaska.gov/static/home/library/pdfs/wildlife/research_pdfs/96_lowry_burkanvov_frost_importance_walleye_pollock_diet_seals.pdf 


1995

Boily, P. (1995). Theoretical heat flux in water and habitat selection of phocid seals and beluga whales during the annual molt. Journal of Theoretical Biology, 172(3), 235–244. https://doi.org/10.1006/jtbi.1995.0020


1994

Frost, K. J., Lowry, L. F., Sinclair, E. H., Ver Hoef, J. M., & McAllister, D. C. (1994). Chapter 6—Impacts on distribution, abundance, and productivity of harbor seals. In Thomas R. Loughlin (ed.): Marine Mammals and the Exxon Valdez (pp. 97–118). Academic Press. https://archive.org/details/marinemammalsexx0000unse

Lowry, L. F., Frost, K. J., Davis, R., Suydam, R. S., & DeMaster, D. P. (1994). Movements and behavior of satellite-tagged spotted seals (Phoca largha) in the Bering and Chukchi Seas. U.S. Department of Commerce NOAA Technical Memo. NMFS-AFSC-38, 71 p. https://apps-afsc.fisheries.noaa.gov/Publications/AFSC-TM/NOAA-TM-AFSC-38.pdf

Lowry, L. F., Frost, K. J., Davis, R., DeMaster, D. P., & Suydam, R. S. (1998). Movements and behavior of satellite-tagged spotted seals (Phoca largha) in the Bering and Chukchi Seas. Polar Biology, 19, 221–230. https://doi.org/10.1007/s003000050238

Lowry, L. F., Frost, K. J., & Pitcher, K. W. (1994). Chapter 12—Observations of oiling of harbor seals in Prince William Sound. In Thomas R. Loughlin (ed.): Marine Mammals and the Exxon Valdez (pp. 209–225). Academic Press. https://archive.org/details/marinemammalsexx0000unse

Weslawski, J. M., Ryg, M., Smith, T. G., & Oritsland, N. A. (1994). Diet of Ringed Seals (Phoca Hispida) in a Fjord of West Svalbard. Arctic, 47(2), 109–114. https://doi.org/10.14430/arctic1279


1990

Kelly, B. P., & Quakenbush, L. T. (1990). Spatiotemporal use of lairs by ringed seals (Phoca hispida). Canadian Journal of Zoology, 68(12), 2503–2512. https://doi.org/10.1139/z90-350


1988

Shults, L. M., & Frost, K. J. (1988). Helminth parasites of ribbon seals, Phoca fasciata, in the Bering Sea and their intermediate hosts. Proc. Helminthol. Soc. Wash., 55(1), 68–73. http://science.peru.edu/COPA/ProcHelmSocWash_V55_N1_1988I.pdf#page=70


1986

Ashwell-Erickson, S., Fay, F. H., Elsner, R., & Wartzok, D. (1986). Metabolic and hormonal correlates of molting and regeneration of pelage in Alaskan harbor and spotted seals (Phoca vitulina and Phoca largha). Canadian Journal of Zoology, 64(5), 1086–1094. https://doi.org/10.1139/z86-163


1984

Bukhtiyarov, Y. A., Frost, K. J., & Lowry, L. F. (1984). New Information on Foods of the Spotted Seal, Phoca largha, in the Bering Sea in Spring. In Francis H. Fay and Gennadii A. Fedoseev (editors), Soviet-American cooperative research on marine mammals (Vols. 1-Pinnipeds, pp. 55–59). https://repository.library.noaa.gov/view/noaa/5600

Lowry, L. F., & Fay, F. H. (1984). Seal eating by walruses in the Bering and Chukchi Seas. Polar Biology, 3, 11–18. https://doi.org/10.1007/BF00265562


1981

Frost, K. J., & Lowry, L. F. (1981). Ringed, Baikal and Capian seals: Phoca hispida Schreber, 1775; Phosa sibirica Gmelin, 1788 and Phoca caspica Gmelin, 1788. In R.J. Harrison (ed.): Handbook of marine mammals, Vol. 2. Seals. (pp. 29–53). Academic Press.

Lowry, L. F., & Frost, K. J. (1981). Feeding and Trophic Relationships of Phocid Seals and Walruses in the Eastern Bering Sea. In D.W. Hood and J.A. Calder, eds. The Eastern Bering Sea Shelf: Oceanography and Resources, Vol. 2. Office of Marine Pollution Assessment, NOAA, BLM. https://www.biodiversitylibrary.org/page/40532600#page/62/mode/1up


1980

Frost, K. J., & Lowry, L. F. (1980). Feeding of ribbon seals (Phoca fasciata) in the Bering Sea in spring. Canadian Journal of Zoology, 58(9), 1601–1607. https://doi.org/10.1139/z80-219

Lowry, L. F., Frost, K. J., & Burns, J. J. (1980a). Feeding of bearded seals in the Bering and Chukchi Seas and trophic interaction with Pacific walruses. Arctic, 33(2), 330–342. https://doi.org/10.14430/arctic2566

Lowry, L. F., Frost, K. J., & Burns, J. J. (1980b). Variability in the diet of ringed seals, Phoca hispida, in Alaska. Canadian Journal of Fisheries and Aquatic Sciences, 37(12), 2254–2261. https://doi.org/10.1139/f80-270


1978

Lowry, L. F., Frost, K. J., & Burns, J. J. (1978). Food of ringed seals and bowhead whales near Point Barrow, Alaska. Canadian Field-Naturalist, 92(1), 67–70. https://doi.org/10.5962/p.346634


1975

Smith, T. G., & Geraci, J. R. (1975). Effect of Contact and Ingestion of Crude Oil on Ringed Seals (Technical Report No. 5; Beaufort Sea Project, p. 66). Canadian Wildlife Service, Department of the Environment. https://www.restco.ca/BSPTR/BSP_TR05_Smith&Geraci.pdf

Stirling, I., Archibald, R., & DeMaster, D. (1975). Distribution and Abundance of Seals in the Eastern Beaufort Sea (Technical Report No. 1; Beaufort Sea Project, p. 58). Canadian Wildlife Service, Department of the Environment. https://www.restco.ca/BSPTR/BSP_TR01_Stirling&Archibald&DeMaster.pdf


1974

Manning, T. H. (1974). Variations in the Skull of the Bearded Seal. University of Alaska. Institute of Arctic Biology. http://hdl.handle.net/11122/1493

1958

McLaren, I. A. (1958). The biology of the ringed seal (Phoca hispida Schreber) in the eastern Canadian Arctic. Bulletin of the Fisheries Research Board of Canada, 118, 1–97. https://waves-vagues.dfo-mpo.gc.ca/library-bibliotheque/40623403_pt3.pdf